The war against the Illumidus Empire, which enslaves and destroys all inhabited planets in its path, has been decisively lost — Earth has fully come under the subjugation of alien conquerors. Harlock and all his comrades have been declared enemies of humanity and expelled from Earth. However, Illumidus did not stop there. Harlock, Tochiro, and Emeralda, under the code names: S – 00999, S – 00998, X – 00001, have been declared wanted. Trying to fight not only against Illumidus but also with many people who wish to receive a reward for the heads of the sought criminals, the sole defenders of Earth continue their journey.