Simon and Kamina grew up in a deep subterranean hamlet, concealed from the legendary surface. Kamina was an impulsive daredevil eager to carve out a reputation, whereas Simon was a shy youngster lacking any clear goals. While digging in the earth one day, Simon uncovered an enigmatic relic that proved to be the ignition key for a long‑forgotten war machine, which they christened Lagann. With this newfound power, the pair repelled an unexpected assault from above, aided by Yoko Littner—a fiery red‑haired warrior armed with a colossal cannon who roams the world above.
After the clash, the sky lay unobstructed before them, and Simon and Kamina set out with Yoko to traverse the desolate surface. They soon joined forces against the Beastmen, humanoid foes that terrorize humanity"s last bastions inside towering mechs known as Gunmen. Despite setbacks, the trio fought valiantly alongside other survivors to reclaim the surface, gradually piecing together a vast interstellar enigma as they confront the looming threat of a cosmic conspiracy while forging unlikely alliances along the way.