The movie follows Anemone, a girl whose father perished in a Tokyo clash seven years before the narrative begins, leaving her clutching only a plush Gulliver and relying on the AI concierge Dominikids for comfort. Today she becomes pivotal in Acid's experimental plan to defeat the seventh Eureka, known as 'Eureka Seven,' a threat that has claimed 2.6 billion lives. As humanity teeters on the edge, every hope rests with Anemone as she plunges into Eureka Seven's core and the fate of all existence hangs in balance for every living being.