In order to safeguard Japan"s tranquility from enigmatic colossal creatures, the Shinkansen Ultra Evolution Institute created the transformable "Shinkalion" robots. Anticipating fresh threats, the institute is now designing a new model named "Shinkalion Z" and accompanying combat‑enhanced units called "Zailiners," which convert ordinary trains into powerful boosters for the Z. Children who exhibit a strong compatibility become pilots of Shinkalion Z, collaborating with institute personnel to battle the re‑emerging titans. The rise of unfamiliar foes—and the meeting of two young heroes—sets the stage for their daring defense. Their synchronized efforts ensure that the nation remains shielded against any looming danger, while their friendship strengthens as they face each new challenge together.